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Satyrinae (browns, satyrs, morpho butterflies, heaths, graylings, marbled whites; order Lepidoptera, family Nymphalidae) Subfamily of small to medium-sized butterflies, which are mostly brown, or orange and black, with eye-spots on the wings. The bases of the wings are often swollen. The butterflies are generally shade-loving. The larvae have a pair of short points at the anal end. Most larvae feed on grasses and sedges. The subfamily is distributed world-wide.