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Eriocraniidae (subclass Pterygota, order Lepidoptera) Family of small, primitive moths with a broad distribution but few species. The venation of the fore wing and hind wing is similar. mandibles are reduced but distinct, and there is a short proboscis. Adults are day fliers. Larvae are leaf-miners and often legless. Pupae have large, curved mandibles, and pupation occurs in the soil in a strong cocoon.