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The Columbia Encyclopedia, Sixth Edition | 2008 | The Columbia Encyclopedia, Sixth Edition. Copyright 2008 Columbia University Press. (Hide copyright information) Copyright

pollen minute grains, usually yellow in color but occasionally white, brown, red, or purple, borne in the anther sac at the tip of the slender filament of the stamen of a flowering plant or in the male cone of a conifer. The pollen grain is actually the male gametophyte generation of seed plants (see reproduction ). Inside the anther, pollen mother cells divide by meiosis to form pollen grains whose nuclei contain half the number of chromosomes characteristic of the parent plant. Each pollen grain contains two sperm nuclei and one tube nucleus. After successful pollination , the pollen germinates on the surface of the stigma of the pistil and produces a tube that grows down through the style to an ovule inside the ovary at the base of the pistil. The sperm nuclei are then discharged into the ovule; one fuses with the egg nucleus (see fertilization ) and the other fuses with the polar nuclei to form endosperm (food-storage tissue) that in many cases nourishes the developing embryo in the seed. This process is basically similar in the conifers, except that in conifers there is no double fertilization and there may be a season's lapse between pollination and fertilization (see cone ). Pollen grains, like sperms, are always produced in much greater quantities than are actually used, particularly by those plants that rely on the wind for pollination (e.g., grasses and conifers). Often clouds of dustlike pollen can be seen floating from wind-pollinated trees. Plants pollinated by insects and birds usually have sticky pollen and conspicuous flowers with colorful petals that often secrete perfume or nectar or both to attract the agents. Although pollen grains are microscopic in size and are thus visible to the human eye only in quantity, they are so diversified in appearance that plants are often identifiable by their pollen alone, e.g., by pathology. The waxy outer covering (which contains proteins and sugar—an additional attraction to pollen-gathering insects) is marked by characteristic patterns of ridges, spines, and knobs and is capable of expanding and contracting in the presence of moisture or dryness. Pollen grains are also remarkable for the length of the tubes some must produce: corn pollen tubes may grow 8 or 10 in. (20.3-25.4 cm) from the stigmas through the filamentous styles (commonly called "silk" ) to the ovaries. The life span of pollen may be less than two hours; its ability to produce the allergic reaction of hay fever continues indefinitely.

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pollen

A Dictionary of Plant Sciences | 1998 | | © A Dictionary of Plant Sciences 1998, originally published by Oxford University Press 1998. (Hide copyright information) Copyright

pollen Collectively, the mass of microspores or pollen grains produced within the anthers of a flowering plant (angiosperm) or the male cones of a gymnosperm.

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MICHAEL ALLABY. "pollen." A Dictionary of Plant Sciences. 1998. Encyclopedia.com. 20 Dec. 2009 <http://www.encyclopedia.com>.

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pollen

A Dictionary of Earth Sciences | 1999 | | © A Dictionary of Earth Sciences 1999, originally published by Oxford University Press 1999. (Hide copyright information) Copyright

pollen Collectively, the mass of microspores (pollen grains) produced by the anthers of a flowering plant (angiosperm) or the male cones of a gymnosperm. Different pollen types are described according to their shapes, apertures, etc. Furrows on the surface of the pollen grain are called ‘sulci’ (sing, sulcus) and monosulcate pollen has a single sulcus. Tricolpate pollens have three, furrow-like, germinal apertures arranged 120° apart and there are many variants of this type.

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AILSA ALLABY and MICHAEL ALLABY. "pollen." A Dictionary of Earth Sciences. 1999. Retrieved December 20, 2009 from Encyclopedia.com: http://www.encyclopedia.com/doc/1O13-pollen.html

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Free newspaper and magazine articles

Free Article Pollen.com Introduces New Features to Help Allergy Sufferers.
Business Wire; 6/2/2008
Free Article Our pollens exceed rest in U.S., making us sneezy and grumpy.(Health)
Newspaper article from: The Register-Guard (Eugene, OR); 4/5/2004
Free Article Pollen Can Hinder Lifting Fingerprints
News Wire article from: AP Online; 4/5/2007

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Pollen.com Introduces New Features to Help Allergy Sufferers.
Business Wire; 6/2/2008; 700+ words ; Pollen History, Pollen Comparison Tools Help Prevent Symptoms PLYMOUTH MEETING, Pa. -- Responding to numerous user requests, Pollen.com, one of the most visited allergy web sites, today announced...
Pollen and Pollination. Special Edition of Plant Systematics & Evolution, Vol. 222
Magazine article from: Kew Bulletin; 1/1/2002; ; 700+ words ; ...Hesse & Ettore Pacini (eds.). Pollen and Pollination. Special Edition of Plant...ISBN 3-211-83514-8 (hardback). `Pollen and Pollination' is a special edition...Its focus, as the title implies, is pollen as a living system from maturity and dispersal...
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Magazine article from: Canadian Journal of Botany; 5/1/2007; ; 700+ words ; ...size and structural characteristics occurs in pollen grains of several species. Pollen grains of Thymus capitatus (L.) Hoffmans...scanning electron microscopy, a quantification of pollen polymorphism of T. capitatus was achieved by...
Pollen can hinder police trying to lift fingerprints
Newspaper article from: Charleston Gazette; 4/7/2007; ; 700+ words ; ATLANTA - All the pollen-covered cars in the South may be an...dust might be a bandit's best friend. Pollen can make it difficult for crime scene...leave behind. In the spring when the pollen is at its worst in the region, police...
Pollen Can Hinder Lifting Fingerprints
News Wire article from: AP Online; 4/5/2007; 700+ words ; ATLANTA - All the pollen-covered cars in the South may be an...dust might be a bandit's best friend. Pollen can make it difficult for crime scene...leave behind. In the spring when the pollen is at its worst in the region, police...
Pollen dispersal models in quaternary plant ecology: assumptions, parameters, and prescriptions.
Magazine article from: The Botanical Review; 1/1/1999; ; 700+ words ; II. Introduction Fossil pollen assemblages are the primary source of...years. Vegetational inferences from pollen assemblages are not straightforward...distortions are introduced by widespread pollen dispersal and by differential pollen...
Pollen forensics may help in fight against terrorism.
M2 Presswire; 4/9/2003; 700+ words ; ...2003-Texas A&M University: Pollen forensics may help in fight against terrorism...LTD RDATE:04082003 COLLEGE STATION - Pollen can be a major irritation to allergy sufferers...University anthropologist Vaughn Bryant. Pollen samples are useful tools that can point...
Our pollens exceed rest in U.S., making us sneezy and grumpy.(Health)
Newspaper article from: The Register-Guard (Eugene, OR); 4/5/2004; 700+ words ; ...heavy loads of tree pollens means more pollen has been hanging around...allergists say. Any grass pollen count over 200 pollens per cubic meter is considered...they surpass 1,000 pollens per cubic meter. Sometimes the pollen here is so thick that...
Pollen's Popping Out All Over; Spring Allergy Season In Full Bloom
Newspaper article from: The Washington Post; 4/16/1996; ; 700+ words ; ...laboratory at Walter Reed, is Washington's official pollen counter. It is her job to catalogue and count these particles -- dozens of species of tree pollen, grass pollen, mold spores and weeds -- the seasonal scourges that...
The Pollen Receptor Kinase LePRK2 Mediates Growth-Promoting Signals and Positively Regulates Pollen Germination and Tube Growth1[W][OA]
Magazine article from: Plant Physiology; 11/1/2008; ; 700+ words ; ...In flowering plants, the process of pollen germination and tube growth is required for successful fertilization. A pollen receptor kinase from tomato (Solanum...been implicated in signaling during pollen germination and tube growth as well...
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