giant clam

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A Dictionary of Earth Sciences

The Columbia Encyclopedia, Sixth Edition

giant clam

The Columbia Encyclopedia, Sixth Edition | 2008 | The Columbia Encyclopedia, Sixth Edition. Copyright 2008 Columbia University Press. (Hide copyright information) Copyright

giant clam common name for the largest bivalve mollusk in the world, Tridacna gigas, also known as the bear's paw clam. The giant clam may weigh over 500 lb (225 kg) and attain a length of over 4 ft (120 cm). The heavy shell is coarsely fluted and toothed. Giant clams are found in the South Pacific and Indian oceans, especially in the Great Barrier Reef. They lie with the hinge downward in the coral reefs, usually in shallow water. The adductor muscles, which cause the shell to close, are a source of food for people of the South Pacific. The shell closes very slowly; stories of human beings trapped within giant clams have never been substantiated. Small giant clam shells have been used as birdbaths and baptismal fonts. An interesting symbiosis occurs between a unicellular green alga ( Zooanthella ) and the clam. The algae live in the tissues of the clam's siphon and mantle; they are able to obtain the sunlight needed for photosynthesis because the clam lies with its valves opening upward and part of the thick, purple mantle extruding over the shell. In addition, there are crystalloid vesicles on the mantle surface that let in sunlight, thus allowing the algae to live deep within the tissues. The clam uses the algae as a supplementary or perhaps even a major source of food. Tridacna gigas is classified in the phylum Mollusca , class Pelecypoda or bivalvia, order Eulamellibranchia, family Tridacnidae.

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hydrothermal vent

A Dictionary of Earth Sciences | 1999 | | © A Dictionary of Earth Sciences 1999, originally published by Oxford University Press 1999. (Hide copyright information) Copyright

hydrothermal vent A place on the ocean floor, on or adjacent to a mid-ocean ridge, from which there issues water that has been heated by contact with molten rock, commonly to about 300°C. The vent water often contains dissolved sulphides. These are oxidized by chemosynthetic bacteria, which fix carbon dioxide and synthesize organic compounds. Near the vents, at temperatures up to 40°C, there are highly productive communities comprising animals that utilize the organic compounds or live symbiotically with the chemosynthetic bacteria; these organisms support carnivores and detritivores. These communities include beard worms (phylum Pogonophora) that completely lack a digestive tract, Munidopsis crabs (superfamily Galatheoidea), giant clams (e.g. Calyptogena magnifica), mussels, acorn worms (class Enteropneusta), and many more. Vent fluids containing high concentrations of iron, manganese, and copper tend to be hot (about 350°C) and black. They are known as ‘black smokers’. ‘White smokers’ flow more slowly, are cooler, and contain high concentrations of arsenic and zinc. See HYDROTHERMAL ACTIVITY, and HYDROTHERMAL MINERAL. See also COLD SEEP.

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